Butterflies of Norway

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COLIAS WERDANDI

English name:	Pale Arctic Clouded Yellow
Norwegian name:	Polargulvinge
Auctor:	(Zetterstedt, 1840)
Original description:	as Colias nastes werdandi in Insecta Lapponica: 908.
Syn.:	- - -
Type locality of nominate species:	Torne Lappland, Sweden.
Worldwide distribution:	Arctic Fennoscandia only; North Norway, Northwest Sweden and
	Northwest Finland, ? Novaya Zelmya Island.
Distribution in Norway:	North Norway; Troms fylke and northern Nordland fylke only.
Subspecies in Norway:	Only the nominate subspecies is known (although the taxon C. nastes zemblida
	Grieshuber, 1998 from Novaya Zemlya Island might be a subspecies of C. werdandi).
Notes:	Until recently, Colias werdandi was regarded to be a subspecies of Colias nastes,
	a species found in North America and Northeast Asia. Some authors, especially
	Russians, have placed C. werdandi as a subspecies of C. tyche (Boeber, 1812),
	a species which inhabits Siberia and Mongolia.
	Recent studies by Jan Haugum, have, however, proved that Colias werdandi is
	morphologically distinct from both C.nastes and C.tyche (ref.: Kudrna, 2002).
	The little know f. christiernssoni Lampa is a rare yellow-orange form
	which has been suggested to be a natural hybrid between C. werdandi
	and C. hecla. This author belives it is just a rare form.
Recognized subspecies:	None.
*Subspecies of C.nastes:*	C.n.nastes Boisduval, 1832 - Greenland, Nunavut, N and NW Canada
	C.n.moina - Churchill
	C.n.thula - North Alaska
	C.n.mongola Alpheraky, 1897 - Altai and Sayan Mts.
	C.n.streckeri Grum-Grshimaïlo, 1895 - N Siberia, Chukot Peninsula
	C.n.dezhnevi Korshunov, 1995 - Magadan Province, Chukotka, Kamchatka
	C.n.flinti Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001 - Wrangal Is. (NE Siberia)
	C.n.magadanica Churkin. Grieshuber, Bogdanov & Zamolodchikov, 2001 - SE Magadan Province.
	C.n.jacutica Kurentzov, 1970 - Eastern Yakutia
	C.n.zemblica Verity, 1911 - Novaya Zemlya Is. (subspecies of nastes, werdandi or tyche?)
Etymology:	Werdandi: Verdandi or Verðandi was one of the three norns, guarding Undarbrunnr, the Weird's Well.
	Verdandi ("Present" or "Necessity"), Urda ("Past" or "Fate") and Skuld ("Future" or "Guilt") weaved the
	thread of fate (Yggdrasil, Norse mythology).

Colias werdandi (males) from North Norway

1 2 3
4 5 6
7 8 9

COLIAS WERDANDI - males

Specimen	EIS	Region	Kommune	Coll. date	Legit	In coll.	Remarks
1	147	TRI	Bardu	VII-1893		ZMO	m
2	127	NSI	Saltdal	09-VII-1918	Johan Rygge	ZMO	m
3	127	NSI	Saltdal	11-VII-1918	Johan Rygge	ZMO	m
4	164	TRI	Nordreisa		Embrik Strand	ZMO	m
5	127	NSI	Saltdal	1918	Johan Rygge	ZMO	m
6	127	NSI	Saltdal	06-VII-1882	Wilhelm Maribo Schøyen	ZMO	m
7	127	NSI	Saltdal	11-VII-1918	Johan Rygge	ZMO	m
8	127	NSI	Saltdal	VII-1918	Johan Rygge	ZMO	m
9	127	NSI	Saltdal	02-VIII-1881	Wilhelm Maribo Schøyen	ZMO	m

Colias werdandi (females) from North Norway

1 2 3
4 5 6
7 8 9

COLIAS WERDANDI - females

Specimen	EIS	Region	Kommune	Coll. date	Legit	In coll.	Remarks
1	127	NSI	Saltdal	12-VII-1918	Johan Rygge	ZMO	f
2	127	NSI	Saltdal	02-VIII-1881	Wilhelm Maribo Schøyen	ZMO	f
3	127	NSI	Saltdal	VII-1918	Johan Rygge	ZMO	f
4	127	TRI	Saltdal	11-VII-1918	Johan Rygge	ZMO	f
5	127	NSI	Saltdal	08-VII-1881	Wilhelm Maribo Schøyen	ZMO	f
6	147	TRI	Bardu	19-VII-1893		ZMO	f
7	155	TRI	Storfjord	14-VII-1894		ZMO	f
8	147	TRI	Bardu	18-VII-1893		ZMO	f
9	127	NSI	Saltdal	11-VII-1918	Johan Rygge	ZMO	f

Colias werdandi from North Norway

1 2
3 4

COLIAS WERDANDI

Specimen	EIS	Region	Kommune	Coll. date	Remarks
1	155	TRI	Balsfjord	08-JUL-1981	m
2	155	TRI	Balsfjord	08-JUL-1981	m
3	155	TRI	Balsfjord	08-JUL-1981	m (ventral view)
4	155	TRI	Balsfjord	03-JUL-1980	f

Habitat of Colias werdandi in North Norway

Habitat of Colias werdandi (Troms fylke, North Norway, elevation 400 m, JUN-2002)

For further notes, photos and maps of this species, check out
the Lepidoptera homepage of the Zoological Museum of Oslo/Norwegian Entomological Society/Skogforsk
(text in Norwegian).